Licorice Compound (glycyrrhizin) Keeps Aging Mice Slim, Strong and Sharp, But the Control Mice Were Obese

Japanese researchers gave middle-aged male ICR mice a low oral dose of glycyrrhizin, the sweet active compound in licorice root, three times a week for twelve months. Treated mice ended the study roughly 19 grams lighter than controls, with no fatty liver at all (controls: 4 of 5), better kidney histology, higher blood Klotho and IGF-1, lower triglycerides, cholesterol, cystatin-C, IL-6 and TNF-alpha, and better performance on activity, grip and rotarod tests. The authors propose that glycyrrhizin blocks the HMGB1 to NF-kB inflammatory loop, which preserves Klotho, which in turn preserves antioxidant defence and organ function. Effect sizes are very large on paper, but the study used five mice per group, no lifespan endpoint, no food intake data, and control animals that reached about 82 grams, which is severe obesity for this strain.

Licorice root has been in the pharmacopoeia of Chinese and Japanese medicine for two thousand years. A group at Suzuka University of Medical Science, working with the licorice manufacturer Cokey Co., has now tested whether its main sweet constituent, glycyrrhizin, can slow the physical decline of ageing mice.

The design was simple. Male ICR mice received either water or 15 mg/kg of glycyrrhizin by oral gavage, three times a week, from eight weeks of age until twelve months. A separate group of ten-week-old mice provided a young reference point.

The result that jumps out first is body weight. Control mice grew to roughly 82 grams. Glycyrrhizin-treated mice stopped at about 63 grams, with the two groups separating from month seven onward. Four of the five controls had fatty liver at autopsy. None of the treated mice did. Kidney tissue told the same story: control kidneys showed enlarged glomeruli, dilated tubules and heavy deposits of advanced glycation end products, and treated kidneys looked closer to young ones.

The behavioural results follow. Old control mice fell off the rotating rod after about nine seconds. Treated mice lasted around forty. Young mice managed the full three minutes, so glycyrrhizin recovered only a slice of what age had taken, but the slice was real. Exploratory activity and forelimb grip strength moved in the same direction.

The blood work is where the authors build their mechanism. Klotho, a kidney-derived protein that suppresses inflammatory signalling and boosts antioxidant enzymes, falls with age. In this study it fell in controls and stayed high in treated mice. IL-6 and TNF-alpha, the two workhorse markers of inflammatory ageing, rose in controls and stayed lower with treatment. Cystatin-C, a kidney function marker, followed the same pattern.

The proposed chain runs from glycyrrhizin blocking HMGB1, an alarm protein released by damaged and senescent cells, through reduced NF-kB signalling, to preserved Klotho, to protected kidneys and muscles.

But, the control mice were not normal old mice, they were extremely fat old mice. Anything that stops a mouse gaining twenty grams of fat will improve its liver, kidneys, inflammatory markers and ability to stay on a rotating rod. Whether glycyrrhizin does anything beyond that is not established here. No mouse in this study was followed to death, so nothing about lifespan was measured. [Confidence: High]

Actionable Insights

The most defensible takeaway is that a modest dose of a licorice compound stopped middle-aged mice becoming obese, and almost everything else in the paper follows from that.

On magnitude: treated mice were 23 percent lighter, had 26 percent lower triglycerides and 22 percent lower cholesterol, 37 percent lower IL-6, 33 percent lower cystatin-C and 21 percent higher blood Klotho. In standardised terms most of these are Cohen’s d values between 2 and 3.3. For context, a d of 0.8 is conventionally called a large effect. Values above 2 in a five-animal-per-group study almost always shrink when repeated, so treat these as an upper bound rather than an expectation.

Scaling the dose by body surface area, 15 mg/kg in a mouse corresponds to roughly 85 mg for a 70 kg adult, three times a week, or about 36 mg a day averaged. That sits under the 100 mg per day of glycyrrhizic acid that European advisory bodies treat as the upper habitual intake.

The safety point matters more than the benefit. Glycyrrhizin inhibits 11-beta-hydroxysteroid dehydrogenase type 2 and can raise blood pressure and drop potassium. Anyone on diuretics, antihypertensives or with existing hypertension should not self-experiment here. Deglycyrrhizinated licorice, the common supplement form, contains none of the compound tested. [Confidence: Medium]

Context and Source

  • Open Access Paper: Anti-aging effects of glycyrrhizin via maintaining Klotho levels
  • Authors: Shiho Ohnishi, Keiichi Hiramoto, Nobuji Yoshikawa, Shosuke Kawanishi
  • Institutions: Faculty of Pharmaceutical Sciences, Suzuka University of Medical Science, Mie, Japan; Matsusaka R&D Center, Cokey Co., Ltd., Mie, Japan
  • Country: Japan
  • Journal: Journal of Clinical Biochemistry and Nutrition, July 2026.
  • Conflict of interest: author NY is an employee of Cokey Co., Ltd., which manufactures glycyrrhizin for medicinal and cosmetic use. The company also supplied the test compound.
  • Impact Score: The impact score of this journal is 1.9 (2025 Journal Impact Factor; SJR 0.608, Q2 in Medicine miscellaneous, Q3 in Clinical Biochemistry and in Nutrition and Dietetics), evaluated against a typical high-end range of 0 to 60+ for top general science, therefore this is a Low impact journal.

Biomarker Data (Effect Size Extraction)

The paper reports no numeric tables and no summary statistics in the text. Every value below was read off the bar charts in Figures 1, 3 and 4, so treat them as approximations with perhaps 5 percent reading error. Cohen’s d is the difference between two group means divided by their pooled standard deviation. A d of 0.2 is small, 0.5 medium, 0.8 large. Values here are extreme because the standard deviations are small and n is 5. Hedges’ g applies a small-sample correction and is the more honest number.

Comparison is 12-month control versus 12-month glycyrrhizin, n = 5 per group.

Endpoint Control (mean, SD) Glycyrrhizin (mean, SD) Relative change Cohen’s d Hedges’ g Percent of the age-related deficit reversed
Body weight (g) 82, 6 63, 4.5 23 percent lower 3.58 3.24 not applicable
Triglyceride (mg/dL) 253, 13 187, 33 26 percent lower 2.63 2.38 35 percent
Cholesterol (mg/dL) 94, 10 73, 9 22 percent lower 2.21 1.99 35 percent
Klotho (pg/mL) 58, 4 70, 4 21 percent higher 3.00 2.71 80 percent
IGF-1 (ng/mL) 10.5, 1.4 15.5, 2.2 48 percent higher 2.71 2.45 64 percent
Cystatin-C (ng/mL) 533, 62 357, 42 33 percent lower 3.32 3.00 70 percent
IL-6 (pg/mL) 58, 6 36.5, 7 37 percent lower 3.30 2.98 49 percent
TNF-alpha (pg/mL) 40, 7 29, 7 28 percent lower 1.57 1.42 44 percent
Activity (counts/15 min) 280, 75 437, 45 56 percent higher 2.54 2.29 53 percent
Grip strength (kgf) 1.34, 0.13 1.59, 0.07 19 percent higher 2.39 2.16 44 percent
Rotarod latency (s) 9, 2 42, 30 367 percent higher 1.55 1.40 19 percent

The IGF-1 problem. The paper treats rising IGF-1 as an anti-ageing signal, on the grounds that Klotho supports the GH/IGF-1 axis and that axis maintains muscle and bone. This runs against the dominant geroscience position, where reduced GH/IGF-1 signalling (Ames and Snell dwarfs, GHRKO, IGF1R heterozygotes) is one of the most reproducible lifespan-extending manipulations in mammals. Klotho’s better-known action is in fact to inhibit insulin and IGF-1 signalling, not to raise it. The paper does not engage with this tension at all. In obese, hyperinsulinaemic middle-aged animals, low IGF-1 may simply be a marker of metabolic derangement rather than a longevity signal, but the authors do not make that argument, and as written the framing is a weak point. [Confidence: High that the tension exists; Medium on the resolution]

Novelty

What is new here, narrowly stated:

  1. First report that chronic low-dose glycyrrhizin preserves circulating Klotho during normal ageing in mice. Prior glycyrrhizin work covered hepatoprotection, colorectal carcinogenesis, diabetic nephropathy and endothelial senescence in disease models, not Klotho in an ageing time course. [Confidence: Medium, this is the authors’ own claim and I have not verified it exhaustively]
  2. First report of glycyrrhizin improving age-related motor function decline (rotarod, grip, open field). The authors state explicitly that no prior reports exist for motor decline, and that appears correct.
  3. A twelve-month continuous dosing time course showing weight divergence beginning at month seven, which is a longer exposure than most glycyrrhizin studies.

What is not new: the HMGB1 to NF-kB mechanism, the Klotho biology, the metabolic effects of glycyrrhizin, and the general observation that reducing inflammatory load improves aged phenotypes. The mechanistic figure is a literature synthesis, not a finding.

Critical Limitations

Ranked by how much they should move your posterior.

  1. The control group is not a normative ageing control. At approximately 82 grams these are morbidly obese mice, against a strain reference nearer 45 to 55 grams. Every downstream result, hepatic steatosis, renal AGE deposition, IL-6, TNF-alpha, cystatin-C, rotarod failure at nine seconds, is a textbook consequence of that obesity. This study is much better described as a diet-induced-obesity-like prevention study than as an ageing study. The correct control for the claim being made would have been a weight-matched pair-fed group. It does not exist. [Confidence: High]
  2. Food intake was never measured. Glycyrrhizin has known effects on taste, gut and metabolism, and gavage three times a week is itself a stressor. Without intake data the entire effect could be a caloric intake difference. This is the single most important missing dataset in the paper. [Confidence: High]

Actionable Intelligence

Compound: glycyrrhizin (glycyrrhizic acid; the study used glycyrrhizic acid monoammonium). Source study: Ohnishi et al., J Clin Biochem Nutr 2026, 79(1): 104-110.

1. The Translational Protocol

Human Equivalent Dose, body surface area normalisation

FDA guidance formula:

HED (mg/kg) = Animal dose (mg/kg) x (Animal Km / Human Km)

Km values from the FDA table: mouse Km = 3, human (60 kg adult) Km = 37.

Step by step:

  • Animal dose in the study: 15 mg/kg, oral gavage, three times per week
  • Km ratio: 3 / 37 = 0.081
  • HED per administration: 15 x 0.081 = 1.22 mg/kg
  • For a 60 kg adult: 1.22 x 60 = 73 mg per dose
  • For a 70 kg adult: 1.22 x 70 = 85 mg per dose
  • For a 80 kg adult: 1.22 x 80 = 97 mg per dose
  • Weekly exposure for a 70 kg adult: 85 x 3 = 256 mg per week
  • Averaged daily exposure: 256 / 7 = 36.5 mg per day, equal to 0.52 mg/kg per day

Cross-check against the human no-effect level. The Scientific Committee on Food identifies a NOAEL of 2 mg/kg body weight per day, roughly 130 mg per person per day, from the Bijlsma and van Gelderen human volunteer work, and sets a provisional upper limit of 100 mg per day of glycyrrhizic acid from all sources.

  • On an averaged daily basis: 0.52 against a NOAEL of 2 mg/kg/day, a margin of about 3.8 fold
  • On a dosing-day basis: 1.22 against 2 mg/kg/day, a margin of about 1.6 fold

That is a thin margin for a chronic protocol, and it is a margin against an endocrine effect that has no therapeutic upside. The 85 mg per dose figure sits just under the 100 mg per day advisory ceiling, so a person taking this three times a week is operating at roughly one third of the regulatory ceiling on average and at 85 percent of it on dosing days.

Salt-form note: the study used glycyrrhizic acid monoammonium (molecular weight about 840) rather than the free acid (about 822). The correction is under 3 percent and does not change any conclusion.

Caveat on the method itself: BSA normalisation is a first-in-human starting-dose heuristic from FDA guidance. It does not account for the species difference that matters most here, which is that rodents are markedly less sensitive to 11-beta-HSD2 inhibition than humans are. The true human-equivalent risk at this dose is higher than the arithmetic suggests.

Guidance for industry: estimating the maximum safe starting dose in initial clinical trials for therapeutics in adult healthy volunteers, FDA (2005), A simple practice guide for dose conversion between animals and human (2016), Opinion of the Scientific Committee on Food on glycyrrhizinic acid and its ammonium salt (2003)

Pharmacokinetics

  • Oral bioavailability of glycyrrhizin as such: poor. Glycyrrhizin is barely absorbed intact from the gastrointestinal tract. It requires hydrolysis by intestinal bacterial beta-glucuronidase to 18-beta-glycyrrhetinic acid, which is then almost completely absorbed. This is the single most important PK fact for anyone reading the mechanism section, because the systemic species is glycyrrhetinic acid, not the HMGB1-binding parent molecule.
  • Absorption of the metabolite: near complete once formed, but formation is microbiome dependent. Gut microbiome perturbation changes glycyrrhizic acid bioavailability in rats.
  • Tmax: highly variable. After a 600 mg oral dose the circulating glucuronide metabolite peaked anywhere from 1.5 to 39 hours later, reflecting the bacterial activation step and enterohepatic recycling.
  • Elimination: slow and multi-peaked. Plasma concentrations show several peaks over roughly 50 hours from enterohepatic recirculation, and complete elimination of glycyrrhetic acid takes several days. Only 0.31 to 0.67 percent is excreted in urine; elimination is predominantly biliary.
  • Practical consequence: a “three times a week” schedule does not produce washout between doses. Effective exposure is closer to continuous than intermittent, which matters because the mineralocorticoid effect is cumulative, and because the intermittent-dosing rationale that makes rapamycin tolerable does not transfer here.
  • Single clean half-life value: not available in the sources retrieved. The literature reports multi-compartment behaviour rather than a single terminal half-life. Treat any single number you see quoted with suspicion.

Intestinal bacterial hydrolysis is indispensable to absorption of 18-beta-glycyrrhetic acid after oral administration of glycyrrhizin in rats (1994), Bioavailability study of glycyrrhetic acid after oral administration of glycyrrhizin in rats (1996), The effect of gut microbiome perturbation on the bioavailability of glycyrrhizic acid in rats (2025), Analysis and pharmacokinetics of glycyrrhizic acid and glycyrrhetinic acid in humans and experimental animals (1994), Hazard assessment of glycyrrhizic acid from liquorice, VKM Report 2018:09

Safety and Toxicity

Acute toxicity (LD50). Oral LD50 in rats is 14.2 g/kg (males) and 18.0 g/kg (females). In mice it exceeds 7.5 g/kg for both sexes. Glycyrrhizate salts in mice span 1,220 to 12,700 mg/kg. Acute toxicity is effectively a non-issue; this compound does not kill by overdose at anything resembling a supplement dose.

Chronic toxicity. A 96-week mouse study at up to 407 mg/kg showed no evidence of chronic toxicity or tumourigenicity. Short-term rat studies showed minor red cell and hepatic enzyme changes with a no-effect level of 0.31 to 0.63 g extract/kg. The organ toxicity profile is benign. The problem is not organ toxicity, it is endocrine.

NOAEL, the number that actually governs use: 2 mg/kg body weight per day in humans, roughly 130 mg per person per day, from human volunteer studies, based on potassium depletion as the endpoint. The SCF’s provisional upper intake limit is 100 mg per day. Pharmacokinetic modelling predicts roughly 4 per 10,000 exposed people will show pseudohyperaldosteronism symptoms at that 100 mg per day limit, and the SCF explicitly flags that subgroups including hypertensives and people with genetic 11-beta-HSD2 variants are not protected by it.

Phase I and controlled human safety data. These exist, and they are informative because they come from a licensed pharmaceutical product rather than from supplement use.

  • A double-blind randomised placebo-controlled phase I/II trial of intravenous glycyrrhizin in chronic hepatitis C was conducted and published in 1999.
  • Stronger Neo-Minophagen C is a licensed intravenous glycyrrhizin preparation in Japan and elsewhere for abnormal hepatic function in chronic liver disease. Each 2 mL ampoule delivers 4 mg glycyrrhizin; the daily regimen is 40 to 60 mL, that is 80 to 120 mg glycyrrhizin intravenously per day, maximum 100 mL, that is 200 mg.
  • Registration-dossier clinical data for that product report hypokalaemia in 7.5 percent and hypertension in 4.4 percent of treated hepatic-disease patients.
  • Listed serious adverse reactions: pseudoaldosteronism with severe hypokalaemia, hypertension, sodium and fluid retention, oedema; hypokalaemic myopathy; rhabdomyolysis (reported with oral preparations); rare shock.
  • Contraindications: aldosteronism, myopathy, hypokalaemia, prior hypersensitivity.

Read that in context. At an intravenous dose roughly in the same range as the calculated HED, in a monitored clinical population, roughly 1 in 13 patients develops hypokalaemia and roughly 1 in 23 develops hypertension. That is your real-world adverse event rate. It is not rare.

Meta-analytic human effect at or above 100 mg per day: systolic blood pressure +5.45 mmHg (95 percent CI 3.51 to 7.39), diastolic +3.19 mmHg (95 percent CI 0.10 to 6.29), plasma potassium −0.33 mmol/L (95 percent CI −0.42 to −0.23), across 18 studies and 337 subjects, with dose-response correlation r squared 0.55 systolic and 0.65 diastolic.

Liver and kidney signals. Hepatically the signal is favourable, not adverse: glycyrrhizin preparations lower transaminases in chronic liver disease and have been used for decades on that basis. Renally the drug does not appear directly nephrotoxic, but the mineralocorticoid effect causes sodium and water retention and potassium wasting, and severe cases produce hypokalaemic nephropathy and acute kidney injury secondary to rhabdomyolysis. The paper’s claim of renal protection in mice must be set against a documented human renal-endocrine liability.

Hazard assessment of glycyrrhizic acid from liquorice, VKM Report 2018:09, Opinion of the Scientific Committee on Food on glycyrrhizinic acid (2003), Intravenous glycyrrhizin for the treatment of chronic hepatitis C: a double-blind, randomized, placebo-controlled phase I/II trial (1999) (publisher returned 403 on direct fetch; record located via live search), Stronger Neo-Minophagen C assessment report, composition, dosing and adverse reactions, The association between consistent licorice ingestion, hypertension and hypokalaemia: a systematic review and meta-analysis (2017), Liquorice toxicity: a comprehensive narrative review (2023), Final report on the safety assessment of glycyrrhetinic acid and glycyrrhizates, CIR (2007)

CYP450 and transporter interactions

  • CYP3A: modest induction, and it is clinically relevant by bioequivalence criteria. In 16 healthy men in a randomised crossover design, 14 days of glycyrrhizin followed by midazolam on day 15 reduced midazolam AUC from 196.4 to 151.3 ng·h/mL, geometric mean ratio 0.77 (90 percent CI 0.70 to 0.89), Cmax ratio 0.83 (90 percent CI 0.74 to 1.01). The authors called it a modest but clinically relevant induction. Translation: expect roughly a 20 percent reduction in exposure to CYP3A4 substrates.
  • P-glycoprotein: no meaningful effect. Continuous glycyrrhizin did not alter the pharmacokinetics of the P-gp probe substrate talinolol in healthy volunteers.
  • Other CYPs: licorice species and individual constituents inhibit several CYPs in vitro, but the in vitro panel does not predict the in vivo human result, which for CYP3A came out as induction, not inhibition. Do not extrapolate from the in vitro papers.
  • Electrolyte-mediated pharmacodynamic interactions (not CYP, and more important): digoxin (hypokalaemia potentiates digoxin toxicity), loop and thiazide diuretics (additive potassium loss), corticosteroids (glycyrrhetinic acid blocks cortisol inactivation, amplifying steroid effect), antihypertensives (opposed), warfarin, and MAO inhibitors.

Effect of glycyrrhizin on the activity of CYP3A enzyme in humans (2010), PubMed record (2010), Lack of effect of continuous glycyrrhizin administration on the pharmacokinetics of the P-glycoprotein substrate talinolol in healthy volunteers (2012), Cytochrome P450 inhibition by three licorice species and fourteen licorice constituents (2018), Licorice, Merck Manual (drug interactions)

2. Biomarker Verification: what actually verifies target engagement

Separate three questions: is the compound in you, is it hitting the target the paper claims, and is it hitting the target it is actually known to hit.

Tier 1, validated, confirms the known pharmacology (use these):

  • Urinary or salivary cortisol to cortisone ratio. This is the definitive target-engagement marker for glycyrrhetinic acid, because 11-beta-HSD2 inhibition is the one human pharmacodynamic effect that is beyond dispute. A rising ratio means the drug is systemically active. It is also, awkwardly, the marker of the adverse mechanism.
  • Serum potassium. Falls with exposure. Cheap, universal, and the most actionable single number.
  • Plasma renin activity and serum aldosterone. Both suppressed. The signature of pseudoaldosteronism is low renin, low aldosterone, elevated cortisol to cortisone ratio, and rising blood pressure. If you see that pattern you have confirmed engagement and you should stop.
  • Home blood pressure, measured properly over 7 days. Detects the 5 mmHg-scale effect that a single clinic reading will not.
  • ALT and AST. The only human-validated efficacy readout for this compound, and only in people who have elevated transaminases to begin with.

Feasibility and ROI

Sourcing. Three tiers, all real.

  • Supplement, over the counter, trivially available. Standardised licorice root extracts at 20 to 30 percent glycyrrhizin are sold widely as capsules, powders and tinctures. Label accuracy for glycyrrhizin content is the main issue; independent testing programmes exist for this category. Getting 85 mg of glycyrrhizin is easy. Getting a known 85 mg is harder.
  • Prescription. Stronger Neo-Minophagen C, intravenous, licensed in Japan and several Asian markets for chronic hepatic dysfunction, at 80 to 120 mg glycyrrhizin per day. Also Glycyron tablets. Not available in the US or EU for this purpose, and there is no reason a healthy person would pursue an intravenous route for a mouse result.
  • Research chemical. Glycyrrhizic acid ammonium salt is a bulk food and pharmaceutical ingredient, sold by the kilogram as a sweetener and flavour, USP and food grade. Cheap and abundant. It is also, relevantly, the manufacturing business of the company that co-authored this paper.

Cost. Negligible on any reading. Licorice extract is among the cheapest botanicals on the market, and glycyrrhizin as a bulk sweetener costs a small fraction of what a month of any real longevity intervention costs. Exact current per-unit pricing: unverified in live search, but the order of magnitude is single-digit to low double-digit dollars per month for the calculated dose, and this figure does not change the analysis.

Cost versus effect, honestly stated. The cost is near zero and so is the expected benefit in a healthy person. What you are buying at 85 mg three times a week is:

  • Expected benefit: nothing demonstrated in humans for ageing, frailty, kidney function or Klotho. The only human-supported efficacy signal is a transaminase reduction in people with existing liver disease, which is not the target population here.
  • Expected cost: an endocrine effect with a Level A human effect size of about +5.4 mmHg systolic and −0.33 mmol/L potassium at doses in this range, and a documented 7.5 percent hypokalaemia and 4.4 percent hypertension incidence in monitored patients at comparable pharmaceutical doses.
  • Marginal gain over things already in a standard stack: zero to negative. Anything that raises systolic pressure by 5 mmHg over years is working directly against the largest single modifiable determinant of cardiovascular and all-cause mortality. The mouse benefits in this paper are almost certainly a weight-gain-prevention effect, and there are many ways to prevent weight gain that do not inhibit 11-beta-HSD2.

ROI verdict: negative. Not because the compound is dangerous at these doses for most people, but because there is no demonstrated human upside to weigh against a small, real, cumulative, Level A downside. [Confidence: High]